A. Klein, B. E. Vaissiere, J. H. Cane, I. Steffan-dewenter, S. A. Cunningham et al., Importance of pollinators in changing landscapes for world crops, Proc. R. Soc. B Biol. Sci, vol.274, pp.303-313, 2007.
URL : https://hal.archives-ouvertes.fr/hal-02669190

D. Goulson, E. Nicholls, C. Botías, and E. L. Rotheray, Bee declines driven by combined stress from parasites, pesticides, and lack of flowers, Science, vol.347, 2015.

L. M. Brutscher, A. J. Mcmenamin, and M. L. Flenniken, The buzz about honey bee viruses, PLoS Pathog, vol.12, 2016.

L. Wilfert, G. Long, H. C. Leggett, P. Schmid-hempel, R. Butlin et al., Deformed wing virus is a recent global epidemic in honey bees driven by Varroa mites, Science, vol.351, pp.594-597, 2016.

G. Di-prisco, F. Pennacchio, E. Caprio, H. F. Boncristiani, J. D. Evans et al., Varroa destructor is an effective vector of Israeli acute paralysis virus in the honey bee, Apis mellifera, J. Gen. Virol, vol.92, pp.151-155, 2011.

T. Takeuchi, R. Takahashi, T. Kiyoshi, M. Nakamura, Y. Minoshima et al., The origin and genetic diversity of the yellow-legged hornet, Vespa velutina introduced in Japan, Insectes Sociaux, vol.3, pp.313-320, 2017.

A. Perrard, J. Haxaire, A. Rortais, and C. Villemant, Observations on the colony activity of the Asian hornet Vespa velutina Lepeletier 1836 (Hymenoptera: Vespidae: Vespinae) in France, Ann. Soc. Entomol. Fr, vol.45, pp.119-127, 2009.

J. K. Kim, M. Choi, and T. Y. Moon, Occurrence of Vespa velutina Lepeletier from Korea, and a revised key for Korean Vespa species (Hymenoptera:Vespidae), Entomol. Res, vol.2, pp.112-115, 2006.

M. B. Choi, S. J. Martin, and J. W. Lee, Distribution, spread, and impact of the invasive hornet Vespa velutina in South Korea, Entomol. Res, vol.15, pp.473-477, 2012.

C. Villemant, D. Zuccon, Q. Rome, F. Muller, G. O. Poinar et al., Can parasites halt the invader? Mermithid nematodes parasitizing the yellow-legged Asian hornet in France, PeerJ, vol.3, p.947, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02557720

J. Haxaire, J. P. Bouguet, and J. P. Tamisier, Vespa velutina Lepeletier, 1836, une redoutable nouveauté pour la faune de France (Hym., Vespidae), Bull. Soc. Entomol. Fr, vol.111, p.194, 2006.

C. Villemant, M. Barbet-massin, A. Perrard, F. Muller, O. Gargominy et al., Predicting the invasion risk by the alien bee-hawking Yellow-legged hornet Vespa velutina nigrithorax across Europe and other continents with niche models, Biol. Conserv, vol.144, pp.2142-2150, 2011.

C. Robinet, E. Darrouzet, and C. Suppo, Spread modelling: A suitable tool to explore the role of human-mediated dispersal in the range expansion of the yellow-legged hornet in Europe, Int. J. Pest Manag, vol.1, pp.1-10, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02627594

Q. Rome, L. Dambrine, C. Onate, F. Muller, C. Villemant et al., Spread of the invasive hornet Vespa velutina Lepeletier, 1836, in Europe in 2012, vol.118, pp.21-22, 2013.

S. Demichelis, A. Manino, G. Minuto, M. Mariotti, and M. Porporato, Social wasp trapping in north west Italy: Comparison of different bait-traps and first detection of Vespa velutina, Bull. Insectol, vol.67, pp.307-317, 2014.

K. Tan, J. Wu, Q. Liu, and D. Y. Zhou, Reduction of foraging activity by A.cerana colonies attacked by Vespa Velutina, J. Bee, vol.2, pp.7-8, 2004.

F. Requier, Q. Rome, G. Chiron, D. Decante, S. Marion et al., Predation of the invasive Asian hornet affects foraging activity and survival probability of honey bees in Western Europe, J. Pest Sci, vol.92, pp.567-578, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02626275

O. Yañez, H. Q. Zheng, F. L. Hu, P. Neumann, and V. Dietemann, A scientific note on Israeli acute paralysis virus infection of Eastern honeybee Apis cerana and vespine predator Vespa velutina, Apidologie, vol.5, pp.587-589, 2012.

M. Garigliany, B. Taminiau, N. El-agrebi, D. Cadar, G. Gilliaux et al., Moku virus in invasive Asian hornets, vol.12, 2016.

M. Mazzei, M. Forzan, G. Cilia, S. Sagona, L. Bortolotti et al., First detection of replicative deformed wing virus (DWV) in Vespa velutina nigrithorax, Bull. Insectol, vol.2, pp.211-216, 2018.

Q. Y. Yue, D. Y. Qiu, Y. W. Huang, and G. X. Liu, Application of DNA barcoding to species identification of an unknown insect larva, Chin. J. Health Lab. Technol, vol.3, pp.615-617, 2011.

M. Matsuura and P. Vespa, The Social Biology of Wasps

K. G. Ross and R. W. Matthews, , p.30, 1991.

D. Bigot, A. Dalmon, B. Roy, C. Hou, M. Germain et al., The discovery of Halictivirus resolves the Sinaivirus phylogeny, J. Gen. Virol, vol.98, pp.2864-2875, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02140702

G. H. Sguazza, F. J. Reynaldi, C. M. Galosi, and M. R. Pecoraro, Simultaneous detection of bee viruses by multiplex PCR, J. Virol. Methods, vol.194, pp.102-106, 2013.

K. Katoh, K. Misawa, K. Kuma, and T. Miyata, MAFFT: A novel method for rapid multiple sequence alignment based on fast Fourier transform, Nucleic Acids Res, vol.30, pp.3059-3066, 2002.

M. Kearse, R. Moir, A. Wilson, S. Stones-havas, M. Cheung et al., Geneious Basic: An integrated and extendable desktop software platform for the organization and analysis of sequence data, Bioinformatics, vol.28, pp.1647-1649, 2012.

S. Guindon, J. F. Dufayard, V. Lefort, M. Anisimova, W. Hordijk et al., New algorithms and methods to estimate maximum-likelihood phylogenies: Assessing the performance of PhyML 3.0, Syst. Biol, vol.3, pp.307-321, 2010.
URL : https://hal.archives-ouvertes.fr/lirmm-00511784

V. Lefort, J. E. Longueville, O. Gascuel, and . Sms, Smart model selection in PhyML, Mol. Biol. Evol, vol.34, pp.2422-2424, 2017.
URL : https://hal.archives-ouvertes.fr/lirmm-01794206

M. Anisimova and O. Gascuel, Approximate likelihood-ratio test for branches: A fast, accurate, and powerful alternative, Syst. Biol, vol.55, pp.539-552, 2006.
URL : https://hal.archives-ouvertes.fr/lirmm-00136658

G. Chen, Z. W. Wang, Y. Qin, and W. B. Sun, Seed dispersal by hornets: An unusual insect-plant mutualism, J. Integr. Plant Biol, vol.59, pp.792-796, 2017.

S. López, M. González, A. Goldarazena, . Vespa, and . Lepeletier, Hymenoptera: Vespidae): First records in Iberian Peninsula, vol.41, pp.439-441, 2011.

H. F. Boncristiani, J. D. Evans, Y. Chen, J. Pettis, C. Murphy et al., In vitro infection of pupae with Israeli acute paralysis virus suggests disturbance of transcriptional homeostasis in honey bees (Apis mellifera), PLoS ONE, vol.9, 2013.

H. Ai, X. Yan, and R. Han, Occurrence and prevalence of seven bee viruses in Apis mellifera and Apis cerana apiaries in China, J. Invertebr. Pathol, vol.1, pp.160-164, 2012.

A. Dalmon, P. Gayral, D. Decante, C. Klopp, D. Bigot et al., Viruses in the invasive hornet Vespa velutina. Viruses, vol.11, 1041.
URL : https://hal.archives-ouvertes.fr/hal-02388796

E. Maori, S. Lavi, R. Mozes-koch, Y. Gantman, Y. Peretz et al., Isolation and characterization of Israeli acute paralysis virus, a dicistrovirus affecting honey bees in Israel: Evidence for diversity due to intra-and inter-species recombination, J. Gen. Virol, vol.88, pp.3428-3438, 2008.

G. Palacios, J. Hui, P. L. Quan, A. Kalkstein, K. S. Honkavuori et al., Genetic analysis of Israel acute paralysis virus: Distinct clusters are circulating in the United States, J. Virol, vol.13, pp.6209-6217, 2008.

Q. Diao, B. Li, H. Zhao, Y. Wu, R. Guo et al., Enhancement of chronic bee paralysis virus levels in honeybees acute exposed to imidacloprid: A Chinese case study, Sci. Total Environ, vol.630, pp.487-494, 2018.

T. B. Clark, A filamentous virus of the honey bee, J. Invertebr. Pathol, vol.32, pp.332-340, 1978.

L. Gauthier, S. Cornman, U. Hartmann, F. Cousserans, J. D. Evans et al., The Apis mellifera filamentous virus genome, vol.7, pp.3798-3815, 2015.

U. Hartmann, E. Forsgren, J. D. Charrière, P. Neumann, and L. Gauthier, Dynamics of Apis mellifera filamentous virus (AmFV) infections in honey bees and relationships with other parasites, Viruses, vol.7, pp.2654-2667, 2015.